Plant growth habits are incredibly diverse, ranging from herbaceous, annuals to long-lived woody trees, each habit type occupying a unique ecological niche and bearing unique evolutionary adaptations to their environments. Thus, the topic of growth habit transitions has drawn interest from a diversity of research areas and several outstanding questions remain concerning the rampant evolutionary transitions across plants: What anatomical changes are associated with habit shifts and how does that effect plant biomechanics? Which evolutionary developmental processes were involved in these transitions? Which climatic variables influence habit transitions across plants? How do fine-scale processes at the cell wall influence organismal form? In this symposium, speakers will interrogate transitions in growth habit from different biological scales–– scaling from the contributions of plant cell wall architecture to gross-stem developmental anatomy, to ecological correlations with habit transitions, to macroevolutionary processes, and finally to the biomechanical differences between habit types. In addition to covering a breath of disciplines which interact with habit transitions, talks will cover plants from distantly related lineages, and speakers will represent a diversity of institutions.